I'd say it is difficult to make an definitive statements, as the authors suggest. Given that humans are borderline, it is possible that was a period of more active competition between males and thus no dominant alpha-male model as seen in apes. Correct me if I am wrong, but chimpanzees are closer evolutionary cousins to humans than gorillas. I'm not an evolutionary biologist so I could be way off here, but isn't it possible that our borderline sized testes are the slow evolutionary trend from oversized testes in the human-chimp ancestor as humans have been more dominated by the alpha male model? Of course I could see the other possibility too. Just wondering.
Chrome
"Recollect me darlin, raise me to your lips, two undernourished egos, four rotating hips"